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Cladonia squamosa (Scop.) Hoffm.


Podetia tall, erect, very variable in shape, usually with one or more erect branches, often forming irregular, perforate cups that continue to proliferate from the margins, patchily corticate and densely covered with small, incised, peeling squamules, soredia absent, basal squamules much incised and fan-like, often in dense clumps; apothecia small, brown, in clusters on tips of short branches. Widespread and locally common in woodland, in heathland and on moorland, on rotting wood, on degraded peat and on boulders.

The usual variant, var. squamosa, fluoresces white under UV light and is negative to usual chemical tests. Var. subsquamosa (Nyl. ex Leight.) Vain. is a probably minor chemical variant, K+ yellow-orange, PD+ orange, and UV negative, said to be possibly more robust, also widespread.

Refs: Smith et al. (2009), 335; Purvis et al. (1992), 210; Nordic Lichen Flora (2013) 5: 72, 106 (photo); Dobson (2005), 144 (photo); Dobson (2011), 151 (photo); Britton (2008) 35 (photo); Jahns (1983), 220-221 (photo); Whelan (2011), 80 (photo); van Herk & Aptroot (2004, 2013), 158-9 (photo); Aptroot et al. (2011), 114 (photo); Krog et al. (1994), 168 (photo); Moberg & Holmåson (1984), 145 (photo); Holien & Tønsberg (2008), 88 (photo); Stenroos et al. (2011), 168 (photo); Wirth (1995), 1: 308, 309 (photo); Wirth et al. (2004), 82 (photo); Wirth, Hauck & Schulz (2013), 1: 384 (photo); Valcárcel et al. (2003), 168-9 (photo); Thomson (1967) 128, plates 19, 20, figs. 89d-g (photos); Hinds & Hinds (2007), 207 (photo); Brodo et al. (2001), 271 (photo); Walewski (2007), 52 (photo); McCune & Geiser (2009), 97 (photo); Lichen Atlas of the British Isles 2: 416, 417 (1996).

FGBI: http://fungi.myspecies.info/all-fungi/cladonia-squamosa

Cladonia squamosa var. squamosa, podetium
Cladonia squamosa var. squamosa, colony
Cladonia squamosa var. squamosa, podetium
Var. squamosa: on shaded boulder in oak (Quercus petraea) forest, Aberfoyle, Stirlingshire, March 2009


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Uploaded July 2010, last updated January 2016