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Cladonia verticillata (Hoffm.) Schaer.
 
(= C. cervicornis (Ach.) Flot. subsp. verticillata (Hoffm.) Ahti)


Podetia cup-like, the cups proliferating from the centre to form tiers, i.e. one cup inside another, soredia absent, basal squamules entire or shallowly lobed, lobes rounded, bluish green when fresh, white beneath, not curled upwards when dry; apothecia small, brown, on the margins of the cups. In heathland, in rock crevices, on drystone walls, etc., widespread but local.

Refs: Van Herk & Aptroot (2003), Bibliotheca Lichenologica 86: 193-203 (photos); Smith et al. (2009), 323 (as subsp. of C. cervicornis); Purvis et al. (1992), 207 (as subsp. of C. cervicornis); Nordic Lichen Flora (2013) 5: 85, 110 (photo); Dobson (2005), 134 (as subsp. of C. cervicornis); Dobson (2011), 142 (as subsp. of C. cervicornis); Jahns (1983), 218-219 (photo, as subsp. of C. cervicornis); van Herk & Aptroot (2004, 2013), 162-163 (photo); Aptroot et al. (2011), 120-1 (photos); Krog et al. (1994), 151 (photo, as C. cervicornis); Stenroos et al. (2011) 184 (photo); Wirth (1995), 1: 311-312, 319 (photo, but might be verticillate variant C. cervicornis, same photo in Wirth et al., 2004, 86, and in Wirth, Hauck & Schulz (2013), 1: 394); Hansen & Anderson (1995) 27 (photo, as subsp. of C. cervicornis); Hinds & Hinds (2007), 216-217 (photo); Brodo et al. (2001), 245-246 (photo, as subsp. of C. cervicornis); Thomson (1967) 97-98, plate 5 (photo); McCune & Geiser (2009), 102 (photo); Lichen Atlas of the British Isles 2: 370 (1996) (as subsp. of C. cervicornis).

Van Herk & Aptroot (2003, see above) consider this a separate species from C. cervicornis and regard the characters of the basal squamules, rather than the extent of the proliferation of the podetia, as diagnostic. Recent British accounts have treated them as subspecies, but there is a general consensus in recent Cladonia literature that they should indeed be regarded as separate species, a view with which I agree and now follow here.

FGBI: http://fungi.myspecies.info/all-fungi/cladonia-cervicornis-subsp-verticillata

Cladonia cervicornis subsp. verticillata
In Breckland lichen heath, near Thetford, Norfolk, in Cladonia community that included C. arbuscula, C. ciliata, C. portentosa,
C. gracilis, C. crispata, C. uncialis, C. furcata, C. ramulosa, C. cervicornis (s.s.), C. foliacea, C. rangiformis, February 2010.
 
Cladonia cervicornis subsp. verticillata
Near Thetford, Norfolk, same colony as above, taken after heavy rain, April 2010.
 
Cladonia cervicornis subsp. verticillata
Montane slope near Ben Lawers, Perthshire, April 2003, a hand-held shot in poor light.


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© A.J. Silverside
Uploaded March 2008 (as C. cervicornis subsp. verticillata), last updated January 2016 (first hosted at www-biol.paisley.ac.uk, December 2006)