<!DOCTYPE HTML PUBLIC "-//W3C//DTD HTML 3.2//EN"> <HTML> <HEAD> <TITLE>Cystocoleus ebeneus (images of British lichens)</TITLE> <META NAME="description" CONTENT="Photograph of Cystocoleus ebeneus (lichen)"> <META NAME="keywords" CONTENT="image, images, photograph, lichen, Cystocoleus"> <META NAME="Author" CONTENT="Alan J. Silverside"> </HEAD> <BODY bgcolor="#000000" background="Icback_dr.jpg" TEXT="#000000" LINK="#3672a4" VLINK="#3672a4"> &nbsp; <center><table border=0 bgcolor="#ffffff" width=1000> <tr><td> <a href="Cyphelium_inquinans.html" target="_top"><img src="previous.gif" border=0 alt="Previous page" align=left></a> <a href="Degelia_atlantica.html" target="_top"><img src="next.gif" border=0 alt="Next page" align=right></a> <blockquote> <h5 align=center><font color="#b22222" face="arial">&nbsp;<br>lastdragon.org</font></h5> <h2 align=center><font color="#b22222">Images of British Lichens</font></h2> <hr width="50%"> <p><br></p> <center> <p><font size=5><b>Cystocoleus ebeneus</b></font><font size=4> (Dyllwyn) Thwaites</font></p> <p><br></p> <table width=840 border=0> <tr><td><div align=justify><font face="arial" size=2>Thallus finely filamentous, like patches of lank, black hair, the hyphae seen under a microscope to be contorted, nodulose, running the length of filaments of the photobiont (algal 'partner'), <a href="http://www.bioref.lastdragon.org/Chlorophyta/Trentepohlia.html">Trentepohlia</a>, and forming tight sheaths around them; apothecia unknown. Widespread but more frequent in upland areas of the north and west, on damp rock faces and on damp trunks.</p> <p>Refs: Smith <i>et al.</i> (2009), 369 (line illustration of hyphae); Purvis <i>et al.</i> (1992), 229 (line illustration of hyphae), 230; Wirth (1995), 1: 367 (line illustration of hyphae), 368 (photo); Wirth, Hauck &amp; Schulz (2013), 1: 444 (photo), 445 (line illustration of hyphae); Stenroos <i>et al</i>. (2011), 199 (photo); Puntillo (1996), plate 15 (photo).</p> <p>This lichen requires microscopic confirmation. <i>Racodium rupestre</i> is macroscopically similar and also uses the orange-pigmented <i>Trentepohlia</i> as its photobiont, but under a microscope, the hyphal cells can be seen to be shorter and less contorted. Very useful, comparative, line illustrations are given in Smith <i>et al.</i>, Purvis <i>et al.</i> and Wirth, as cited above. Regrettably the line illustration of <i>C.&nbsp;ebeneus</i> in Dobson (2005, 2011) does not portray a normal state of this species, showing neither the normal orientation of the hyphae along the length of the photobiont filament nor their contorted appearance. In my view and in my experience of demonstrating the lichen to others, the diagram causes confusion.</p> <p><i>Ephebe lanata</i> (page pending) is also very similar, but coarser. It uses the cyanobacterium (blue-green alga) <i>Stigonema</i> as its photobiont. Free-living <i>Stigonema</i> itself resembles <i>Cystocoleus</i> macroscopically, but under the microscope, <i>Stigonema</i> filaments (trichomes) are non-lichenised, lack orange pigment, and the cells are subspherical and commonly in more than one row.</div></font></td></tr> </table> </center></blockquote><center> <table> <tr><td align=center><img src="Cystocoleus_ebeneus_P5080100.jpg" border=1 vspace=5 alt="Cystocoleus ebeneus"></td></tr> <tr><td align=center><font face="arial" size=2>On damp rockface, Ty Canol, Pembrokeshire, May 2009<br>&nbsp;</font></td></tr> <tr><td align=center><img src="Cystocoleus_ebeneus_DSCN1931.jpg" border=1 vspace=5 alt="Cystocoleus ebeneus, photomicrograph"></td></tr> <tr><td align=center><img src="Cystocoleus_ebeneus_DSCN1951.jpg" border=1 vspace=5 alt="Cystocoleus ebeneus, photomicrograph"></td></tr> <tr><td align=center><img src="Cystocoleus_ebeneus_DSCN1963.jpg" border=1 vspace=5 alt="Cystocoleus ebeneus, photomicrograph"></td></tr> <tr><td align=center><img src="Cystocoleus_ebeneus_DSCN1976.jpg" border=1 vspace=5 alt="Cystocoleus ebeneus, photomicrograph"></td></tr> <tr><td align=center><font face="arial" size=2>Photomicrographs of <i>Cystocoleus ebeneus</i>, from Ty Canol thalli shown above, showing hyphae wrapped along <i>Trentepohlia</i> filaments.<br>&nbsp;</font></td></tr> <tr><td align=center><img src="Cystocoleus_ebeneus_IMG_2933.jpg" border=1 vspace=5 alt="Cystocoleus ebeneus"></td></tr> <tr><td align=center><font face="arial" size=2>On damp rockface, Isle of Muck, June 2012 (BLS meeting)</font></td></tr> </table> </center><blockquote><center> <p><br></p> <table border=0> <tr><td><a href="Cyphelium_inquinans.html" target="_top"><img src="baru6.gif" border=0 alt="Previous page"></a></td> <td width="50">&nbsp;</td> <td><a href="Degelia_atlantica.html" target="_top"><img src="baru8.gif" border=0 alt="Next page"></a></td></tr> </table> </center> <p><br></p> <p><hr></p> <center><font face="Arial" size=2> <p><b><a href="index.html" target="_top">Lichens HOME PAGE</a> | <a href="indexP.html" target="_top">Index of Species</a> | <a href="http://www.lastdragon.org/conditions.html" target="_top">Conditions of Use and e-mail contact</a> | <a href="references.html" target="_top">References</a> | <a href="http://www.lastdragon.org/home.html" target="_top">Site Home Page</a></b></p> </font></center> <p><br></p> <font face="Arial" size=2> <p>&copy; A.J. Silverside<br> Uploaded July 2012, last updated September 2013</p> </font> </blockquote> </td></tr> </table></center> </BODY> </HTML>